Amaranthus tortuosus Hornem.

First published in Hort. Bot. Hafn., Suppl.: 107 (1819)
This species is accepted
The native range of this species is Mexico to Tropical America. It is an annual and grows primarily in the seasonally dry tropical biome. It is used as animal food and a medicine, has environmental uses and for food.

Descriptions

Distribution
Biogeografic region: Amazonia, Andean, Guiana Shield, Caribbean, Orinoquia, Pacific. Elevation range: 0–2000 m a.s.l. Native to Colombia. Colombian departments: Amazonas, Antioquia, Arauca, Atlántico, Bolívar, Boyacá, Caldas, Casanare, Cauca, Cesar, Chocó, Córdoba, Cundinamarca, Huila, La Guajira, Magdalena, Meta, Nariño, Norte de Santander, Quindío, Risaralda, San Andrés y Providencia, Santander, Sucre, Tolima, Valle del Cauca.
Habit
Herb.
Ecology
Habitat according IUCN Habitats Classification: forest and woodland, savanna, shrubland, native grassland, wetlands (inland), artificial - terrestrial.
[UPFC]

Bernal, R., G. Galeano, A. Rodríguez, H. Sarmiento y M. Gutiérrez. 2017. Nombres Comunes de las Plantas de Colombia. http://www.biovirtual.unal.edu.co/nombrescomunes/

Vernacular
amaranto, amaranto blanco, bledo, bledo blanco, bledo chico, bledo colorado, bledo colorao, bledo de puerco, bledo hembra, bledo liso, bledo maleza, bledo rojizo, bleo, bleo de puerco, gleo, lirgua, yuyo hembra
[UNAL]

Extinction risk predictions for the world's flowering plants to support their conservation (2024). Bachman, S.P., Brown, M.J.M., Leão, T.C.C., Lughadha, E.N., Walker, B.E. https://nph.onlinelibrary.wiley.com/doi/full/10.1111/nph.19592

Conservation
Predicted extinction risk: not threatened. Confidence: confident
[AERP]

Amaranthaceae, C. C. Townsend. Flora Zambesiaca 9:1. 1988

Morphology General Habit
Erect annual herb, mostly up to c. 90 cm. (rarely to 1.5 m.) tall.
Morphology Stem
Stem rather slender to stout, usually branched, angular, glabrous or increasingly furnished upwards (especially in the inflorescence) with short to rather long, multicellular hairs.
Morphology Leaves
Leaves glabrous, or thinly and shortly pilose on the inferior surface of the primary venation, long-petiolate (petioles up to c. 8.5 cm. long, sometimes longer than the lamina), lamina ovate or rhomboid-ovate, 1.5–8 (12) × 0.7–5 (8) cm., blunt or retuse at the apex with a distinct, fine mucro formed by the percurrent nerve, cuneate (usually shortly so) at the base; leaf axils without spines.
Morphology Reproductive morphology Flowers
Flowers green, in the lower part of the plant in axillary clusters 4–10 cm. in diam., towards the ends of the stem and branches the leafless clusters approximated to form simple or (the terminal at least) branched spikes c. 3–15 (25) cm. long and 6–8 (10) mm. wide. Lower clusters of flowers entirely female, the spikes generally showing a few male flowers at the apices only (rarely in more than the apical 1 cm.), occasionally with male flowers also scattered among the lower female flowers.
Morphology Reproductive morphology Inflorescences Bracts
Bracts and bracteoles deltoid-ovate, pale-membranous with an erect reddish awn formed by the excurrent green midrib, bracteoles somewhat shorter than or subequalling the perianth, rarely slightly exceeding it.
Morphology Reproductive morphology Flowers Perianth
Perianth segments (4) 5, those of the female flowers c. 1.5–2.75 mm. long, narrowly oblong or spathulate oblong, obtuse or sometimes (particularly those approaching the male flowers) acute, mucronulate, frequently with a greenish dorsal vitta above; those of the male flowers broadly lanceolate or lanceolate-oblong, generally acuminate, only the thin midrib green.
Morphology Reproductive morphology Flowers Gynoecium Stigma
Stigmas 3, flexuose or reflexed, c. 0.75–1 mm. long.
Morphology Reproductive morphology Fruits
Capsule subequalling the perianth, ovoid-urceolate, with a short inflated beak below the style base, c. 1.5–1.75 mm., circumcissile, the lid strongly rugulose below the neck.
Morphology Reproductive morphology Seeds
Seed 1–1.25 mm., compressed, black, shining, faintly reticulate.
[FZ]

M. Thulin et al. Flora of Somalia Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS

Morphology General Habit
Erect annual, mostly up to 90(–150) cm, similar in habit to A. spinosus and with similar leaf-size and shape, but often less coarse
Morphology General Spines
Paired axillary spines absent
Morphology Reproductive morphology Inflorescences
Lower flower clusters axillary, female, 4–10 mm in diam.; upper clusters leafless, forming simple or (the terminal at least) branched spikes 3–15(–25) cm long and 6–8(–10) cm wide, the spikes male for up to c. the apical 1 cm
Morphology Reproductive morphology Flowers Tepal
Tepals of female flowers (4–)5, 1.5–2.75 mm, oblong or spathulate-oblong, obtuse or sometimes (especially towards the male flowers) acute, mucronulate, usually with a greenish dorsal vitta above
Morphology Reproductive morphology Flowers Gynoecium Stigma
Stigmas 3
Morphology Reproductive morphology Fruits
Capsule ovoid-urceolate, c. 1.5–1.75 mm, with a short inflated beak below the style-base, circumscissile, the lid strongly rugulose
Morphology Reproductive morphology Seeds
Seeds faintly reticulate, lenticular.
Distribution
S1, 2 of tropical American origin, now widespread in the tropics of both Old and New Worlds.
Note
A. dubius is the only known polyploid Amaranthus, but its origin remains uncertain. Hybrids are said to be frequent where it occurs in association with A. spinosus; the two species are very similar apart from the axillary spines of the latter. Srivasta, Pal & Nair in Rev. Palaeobot. Palynol. 23: 287–291 (1977) claim that the pollen of A. dubius has larger pores than that of A. spinosus, and that the hybrids are distinguishable by the presence of micrograins among the pollen.
[FSOM]

Distribution
Found in Boyacá, Colombia.
Vernacular
Amaranto, Atakco, Bledo, Bledo blanco, Bledo de espina, Bledo rojizo, Bleo, Bleo de puerco, Calalú, Pira, Yuyo
[UPFC]

The Useful Plants of Boyacá project

Ecology
Alt. 0 - 2000 m.
Morphology General Habit
Herb.
Distribution
Native from Colombia.
Vernacular
Atakco.
[UPB]

George R. Proctor (2012). Flora of the Cayman Isands (Second Edition). Royal Botanic Gardens, Kew

Morphology General Habit
Erect herb up to 1 m tall, more or less branched, glabrous or pubescent above
Morphology Leaves
Leaves rhombic-ovate on long, often red petioles, mostly 2–10 cm long, the apex usually acute and mucronate
Morphology Reproductive morphology Flowers
Flowers greenish, in terminal panicles and dense axillary clusters; perianth of 5 segments; stamens 5
Morphology Reproductive morphology Fruits
Utricle with circumscissile dehiscence.
Distribution
Grand Cayman, Cayman Brac. West Indies and continental tropical America, also in tropical Africa.
Ecology
A weed of waste ground.
Note
The weedy, pantropical ‘spiny salalu’ or ‘macca calalu’, Amaranthus spinosus L., has not been recorded from the Cayman Islands, but is to be expected.
[Cayman]

Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

Distribution
Nativa en Colombia; Alt. 0 - 2000 m.; Amazonia, Andes, Guayana y Serranía de La Macarena, Islas Caribeñas, Llanura del Caribe, Orinoquia, Pacífico, Sierra Nevada de Santa Marta, Valle del Cauca, Valle del Magdalena.
Morphology General Habit
Hierba
Conservation
No Evaluada
[CPLC]

Uses

Use Animal Food
Used as animal food.
Use Environmental
Environmental uses.
Use Gene Sources
Used as gene sources.
Use Food
Used for food.
Use Materials
Used as material.
Use Medicines
Medical uses.
[UPFC]

Use Animal Food
Used to feed pigs when food is scarce (Correa & Bernal 1989).
Use Environmental Pollution Control
Used for phytoremediation of chromium, mercury, arsenic, lead, copper and nickel; bioaccumulator (Mellen 2008).
Use Food
Sold in the markets of Rio de Janeiro (Correa & Bernal 1989).
Use Food Vegetables
Leaves - Used in salads (Correa & Bernal 1989).
Use Medicines Genitourinary System Disorders
Used as a diuretic (Correa & Bernal 1989).
Use Medicines Infections & Infestations
Used in the treatment of fever (Correa & Bernal 1989).
Use Medicines Inflammation
Used in poultices in the treatment of inflamed open wounds (Correa & Bernal 1989).
Use Medicines Skin or Subcutaneous Cellular Tissue Disorders
External use in the treatment of allergic skin problems (Correa & Bernal 1989).
Use Medicines Unspecified Medicinal Disorders
Medicinal (State of the World's Plants 2016, Instituto Humboldt 2014).
[UPB]

Common Names

English
Indian Spinach, Marog, wild spinach
Spanish
Bledo, bledo blanco, calalú, curucu, yuyo, amaranto, pira.

Sources

  • Angiosperm Extinction Risk Predictions v1

    • Angiosperm Threat Predictions
    • http://creativecommons.org/licenses/by/4.0
  • Catálogo de Plantas y Líquenes de Colombia

    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Colombian resources for Plants made Accessible

    • ColPlantA 2021. Published on the Internet at http://colplanta.org
    • https://creativecommons.org/licenses/by/4.0/
  • Flora Zambesiaca

    • Flora Zambesiaca
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Flora of Somalia

    • Flora of Somalia
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Flora of the Cayman Islands

    • Flora of the Cayman Islands
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Herbarium Catalogue Specimens

    • Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/
  • Kew Backbone Distributions

    • The International Plant Names Index and World Checklist of Vascular Plants 2024. Published on the Internet at http://www.ipni.org and https://powo.science.kew.org/
    • © Copyright 2023 World Checklist of Vascular Plants. http://creativecommons.org/licenses/by/3.0
  • Kew Living Collection Database

    • Common Names from Plants and People Africa http://www.plantsandpeopleafrica.com/
  • Kew Names and Taxonomic Backbone

    • The International Plant Names Index and World Checklist of Vascular Plants 2024. Published on the Internet at http://www.ipni.org and https://powo.science.kew.org/
    • © Copyright 2023 International Plant Names Index and World Checklist of Vascular Plants. http://creativecommons.org/licenses/by/3.0
  • Kew Science Photographs

    • Copyright applied to individual images
  • Universidad Nacional de Colombia

    • ColPlantA database
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Useful Plants and Fungi of Colombia

    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Useful Plants of Boyacá Project

    • ColPlantA database
    • http://creativecommons.org/licenses/by-nc-sa/3.0
    • http://creativecommons.org/licenses/by/3.0/